John (Zhen-Xiang) Lu, Ph.D.

Image John Lu
Research Scientist

Dr. Lu’s research program is mainly focused on the improvement of crop nutritional composition and food quality to develop novel and value-added bioproducts.

Current research and/or projects

  • Enhancement of protein and resistant starch contents in field pea to promote its uses in functional food application

  • Development of genetically improved field pea varieties and evaluation of flavor, physicochemical and functional characteristics in high-protein pea lines

  • Novel extraction of oil from canola seeds and evaluation of antioxidants on the quality and shelf life of canola oil for potato deep frying

Research and/or project statements


  • Bioproducts and Bioprocessing
  • Starch and Dietary Fibre in Crops
  • Proteins and Amino Acids in Crops

  • Recent Publications

    Liyanagea, D.W.K., Yevtushenko, D.P., Konschuh, M., Bizimungu, B., Lu, Z.X. 2020. Processing strategies to decrease acrylamide formation, reducing sugars and free asparagine content in potato chips from three commercial cultivars. Food Control 119.

    Li, X., Shi, J., Scanlon, M., Xue, S.J., Lu, Z.X. 2020. Effects of pretreatments on physicochemical and structural properties of proteins isolated from canola seeds after oil extraction by supercritical-CO2 process. LWT - Food Science and Technology 110415. doi: 10.1016/j.lwt.2020.110415.

    Lu, Z.X., He, J.F., Zhang, Y.C., Bing, D.J. 2019. Composition, physicochemical properties of pea protein and its application in functional foods. Critical Reviews in Food Science and Nutrition. doi: 10.1080/10408398.2019.1651248.

    Xiong, M., Zhao, M.L., Lu, Z.X., Balasubramanian, P. 2019. Genotypic variation for phenolic compounds in developing and whole seeds, and storage conditions influence visual seed quality of yellow dry bean genotypes. Canadian Journal of Plant Science 100: 284-295.

    He, J.F., Fang, Y.Y. Lu, Z.X., Wang, L.W., Zhao, X.Q., Fu, X.F., Zhao, J., Liu, H.K. 2019. Identification of differentially expressed genes of Haloxylon ammodendron in response to salinity stress. Biologia Plantarum 63: 610-617. doi: 10.32615/bp.2019.066.

    Zhang, Y.C., Kahl, D.H.W, Bizimungu, B., Lu, Z.X. 2018. Effects of blanching treatments on acrylamide, asparagine, reducing sugars and colour in potato chips. Journal of Food Science and Technology 55: 4028-4041.

    Zhou, Y.J., Yang, Q., Zhong, X.J., Tang, H.P., Deng, M., Ma, J., Qi, P.F., Wang, J.R., Chen, G.Y., Liu, Y.X., Lu, Z.X., Li, W., Lan, X.J., Wei, Y.M., Zheng, Y.L., Jiang, Q.T. 2017. Alternative splicing results in a lack of starch synthase IIa-D in Chinese wheat landrace. Genome 61: 201-208. doi: 10.1139/gen-2017-0246.

    Zhang, X.W., Li, S.Y., Zhang, L.L., Yang, Q., Jiang, Q.T., Lu, Z.X., Liu, C.J., Wei, Y.M., Zheng, Y.L. 2016. Structure and expression analysis of genes encoding ADP-glucose pyrophosphorylase large subunit in wheat and its relatives. Genome 59: 501-507.

    Shen, S.A., Hou, H.W., Ding, C.B., Bing, D.J., Lu, Z.X. 2016. Protein content correlates with starch morphology, composition and physicochemical properties in field peas. Can. J. Plant Sci. 96: 404-412.

    Zheng, K., Jiang, Q.T., Wei, L., Zhang, X., Ma, J., Chen, G.Y., Wei, Y.M., Jennifer, M.F., Lu, Z.X., Zheng, Y.L. 2015. Characterization of starch morphology, composition, physicochemical properties and gene expressions in oat. Journal of Integrative Agriculture 14: 20-28.

    Zheng, K., Jiang, Q.T., Zhang, X., Lan, X.J., Dai, S.F., Wei, Y.M., Laroche, A., Lu, Z.X., Zheng, Y.L. 2015. Differences in starch composition and physiochemical properties are influenced by granule types in wheat and its relatives. Pakistan Journal of Botany 46: 2239-2248.


Education and awards

Ph.D. Genetics, Clemson University, U.S.A. 1997.                                           

M.Sc. Plant Genetics & Breeding, Nanjing Agricultural University, P.R.China. 1989.

B.Sc. Horticuture, Nanjing Agricultural University, P.R.China. 1982.                 


Key publications

  1. Wei, L., Wang, Q., Zhang, L.L., Ma, J., Wang, J.R., Qi, P.F., Li, W., Chen, G.Y., Lan, X.J., Deng, M., Lu, Z.X., Wei, Y.M., Zheng, Y.L., Jiang, Q.T. (2015). Genetic analyses of Glu-1S sh in wheat/Aegilops sharonensis hybrid progenies and development of alien HMW-GSs gene-specific markers, 35(12), 1-10.

    2015 - View publication details

  2. Zhao, S., Jiang, Q.T., Ma, J., Wang, J.R., Liu, Y.X., Chen, G.Y., Qi, P.F., Pu, Z.E., Lu, Z.X., Zheng, Y.L., Wei, Y.M. (2015). Characterization and expression analysis of WOX2 homeodomain transcription factor in Aegilops tauschii, 38(1), 79-85.

    2015 - View publication details

  3. Zheng, K., Jiang, Q.t., Wei, L., Zhang, X.w., Ma, J., Chen, G.y., Wei, Y.m., Jennifer, M.F., Lu, Z.x., Zheng, Y.l. (2015). Characterization of starch morphology, composition, physicochemical properties and gene expressions in oat, 14(1), 20-28.

    2015 - View publication details

  4. Wang, X.M., Gaudet, D.A., Liu, W., Frick, M., Puchalski, B., Lu, Z.X., Leggett, F., Kang, Z.S., Laroche, A. (2014). Defence responses including hypersensitive cell death, oxidative burst and defence gene expression in Moro wheat inoculated with Puccinia striiformis, 36(2), 202-215.

    2014 - View publication details

  5. Zhao, S., Jiang, Q.T., Ma, J., Zhang, X.W., Zhao, Q.Z., Wang, X.Y., Wang, C.S., Cao, X., Lu, Z.X., Zheng, Y.L., Wei, Y.M. (2014). Characterization and expression analysis of WOX5 genes from wheat and its relatives, 537(1), 63-69.

    2014 - View publication details

  6. Zhao, Q.Z., Jiang, Q.T., Yang, Q., Ma, J., Wang, J.R., Chen, G.Y., Liu, Y.X., Lu, Z.X., Wei, Y.M., Zheng, Y.L. (2014). Characterization of intergeneric hybrid between common wheat and aegilops sharonensis (Eig) and transfer of alien high molecular weight glutenin subunits into wheat, 42(4), 640-647.

    2014 - View publication details

  7. Jiang, Q.T., Zhao, Q.Z., Yang, Q., Ma, J., Zhang, X.W., Wang, C.S., Wang, X.Y., Cao, X., Lu, Z.X., Wei, Y.M., Zheng, Y.L. (2014). Amphidiploids between tetraploid wheat and Aegilops sharonensis Eig exhibit variations in high-molecular-weight glutenin subunits, 61(2), 299-305.

    2014 - View publication details

  8. Zheng, K., Xu, J., Jiang, Q., Laroche, A., Wei, Y., Zheng, Y., Lu, Z. (2013). Isolation and characterization of an isoamylase gene from rye, 1(2), 127-133.

    2013 - View publication details

  9. Ma, J., Jiang, Q.T., Zhang, X.W., Lan, X.J., Pu, Z.E., Wei, Y.M., Liu, C., Lu, Z.X., Zheng, Y.L. (2013). Structure and expression of barley starch phosphorylase genes, 238(6), 1081-1093.

    2013 - View publication details

  10. He, J.F., Goyal, R., Laroche, A., Zhao, M.L., Lu, Z.X. (2012). Water stress during grain development affects starch synthesis, composition and physicochemical properties in triticale, 56(3), 552-560.

    2012 - View publication details

  11. Jiang, Q.T., Ma, J., Zhao, S., Zhao, Q.Z., Lan, X.J., Dai, S.F., Lu, Z.X., Zheng, Y.L., Wei, Y.M. (2012). Characterization of HMW-GSs and their gene inaction in tetraploid wheat, 140(7-9), 325-335.

    2012 - View publication details

  12. Li, C.Y., Li, C., Lu, Z.X., Li, W.H., Cao, L.P. (2012). Morphological changes of starch granules during grain filling and seed germination in wheat, 64(2), 166-170.

    2012 - View publication details

  13. Jiang, Q.T., Liu, T., Ma, J., Wei, Y.M., Lu, Z.X., Lan, X.J., Dai, S.F., Zheng, Y.L. (2011). Characterization of barley Prp1 gene and its expression during seed development and under abiotic stress, 139(10), 1283-1292.

    2011 - View publication details

  14. Li, C.Y., Li, W.H., Lee, B., Laroche, A., Cao, L.P., Lu, Z.X. (2011). Morphological characterization of triticale starch granules during endosperm development and seed germination, 91(1), 57-67.

    2011 - View publication details

  15. Gaudet, D.A., Wang, Y., Penniket, C., Lu, Z.X., Bakkeren, G., Laroche, A. (2010). Morphological and molecular analyses of host and nonhost interactions involving barley and wheat and the covered smut Pathogen Ustilago hordei, 23(12), 1619-1634.

    2010 - View publication details

  16. Li, C.Y., Li, W.H., Li, C., Gaudet, D.A., Laroche, A., Cao, L.P., Lu, Z.X. (2010). Starch Synthesis and Programmed Cell Death during Endosperm Development in Triticale (×Triticosecale Wittmack), 52(7), 602-615.

    2010 - View publication details

  17. Jiang, Q.T., Wei, Y.M., Lu, Z.X., Liu, T., Wang, J.R., Pu, Z.E., Lan, X.J., Zhen, Y.L. (2010). Characterization of a novel variant HMW-glutenin gene from Elymus canadensis, 32(4), 361-367.

    2010 - View publication details

  18. Jiang, Q.T., Wei, Y.M., Lu, Z.X., Pu, Z.E., Lan, X.J., Zheng, Y.L. (2010). Structural variation and evolutionary relationship of novel HMW glutenin subunits from Elymus glaucus, 147(3), 136-141.

    2010 - View publication details

  19. Xu, J., Frick, M., Laroche, A., Ni, Z.F., Li, B.Y., Lu, Z.X. (2009). Isolation and characterization of the rye Waxy gene, 52(7), 658-664.

    2009 - View publication details

  20. Sun, J.Y., Gaudet, D.A., Lu, Z.X., Frick, M., Puchalski, B., Laroche, A. (2008). Characterization and antifungal properties of wheat nonspecific lipid transfer proteins, 21(3), 346-360.

    2008 - View publication details

Research facility

5403 1st Avenue South
Lethbridge, AB T1J 4B1




Research Scientist at Lethbridge Research and Development Centre, AAFC, Alberta, Canada

Adjunct Professor at University of Lethbridge, Alberta, Canada